It's the one sitting on the gasket that you can't quite make out. Mentioned in some books (e.g. Haynes) as a white O ring, however, they all seem to be black now.
I'd make that a £2000 o-ring.... price for new big ends and mains and everything else you might replace at the same time.
I once fitted a white o-ring (new) that was ø3.5mm section - I thought it was the correct one, being in my oil filter spares box and being the absolutely correct major dimensions. After 20 miles I realised I had an oil leak - a serious one! I shut down and got recovered home. I bought a new oil filter change kit and found the o-ring was ø4mm in section. No leaks after that. I measure them all now, whatever colour they are!